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While Columnaris being only aerobic (preferring high oxygen environments) can occur in tanks with good filtration/circulation, however it is common in a tank with poor Redox/mineralization, overcrowding, high temperatures, and stress (such as an aggressive fish tank where many inhabitants are constantly bullying others).This is NOT to say that sudden shifts in parameters such as spikes in ammonia (assuming an aerobic environment), can will not trigger Columnaris, as this too is a stressor that can allow for an opportunistic Columnaris infection.